1. Hans Spemann - Biography of Freiburgim-Breisgau, in succession to hans Doflein, a spemann's name will alwaysbe associated with his work development, he was awarded the nobel Prize in http://www.nobel.se/medicine/laureates/1935/spemann-bio.html

Hans Spemann was born on June 27, 1869, at Stuttgart. He was the eldest son of the publisher, Wilhelm Spemann. From 1878 until 1888 he went to the Eberhard-Ludwig School at Stuttgart and when he left school in 1888 he spent a year in his father's publishing business. From 1889-1890 he did his military service and then, after a period as a retail bookseller, he entered, in 1891, the University of Heidelberg . There, until he took his preliminary examination in 1893, he studied medicine, and was especially attracted by the work of the comparative anatomist there, Carl Gegenbaur. During the winter of 1893-1894 he studied at the University of Munich , where he became more closely acquainted with August Pauly - a fact of great importance to him. From the spring of 1894 to the end of 1908, he worked in the Zoological Institute at the . In 1895 he took his degree in zoology, botany, and physics (subjects to serve his anatomical studies), having worked under Theodor Boveri, Julius Sachs, and

2. Medicine 1935 b.1869 d.1941. The nobel Prize in Physiology or Medicine 1935 PresentationSpeech hans spemann Biography nobel Lecture. 1934, 1936. http://www.nobel.se/medicine/laureates/1935/

The Nobel Prize in Physiology or Medicine 1935 "for his discovery of the organizer effect in embryonic development" Hans Spemann Germany University of Freiburg im Breisgau Breisgau, Germany b.1869 d.1941 The Nobel Prize in Physiology or Medicine 1935 Presentation Speech Hans Spemann Biography ... Nobel Lecture The 1935 Prize in: Physics Chemistry Physiology or Medicine Literature ... Peace Find a Laureate: Last modified June 16, 2000 The Official Web Site of The Nobel Foundation

3. Hans Spemann Winner Of The 1935 Nobel Prize In Medicine hans spemann, a nobel Prize Laureate in Physiology and Medicine, at the nobelPrize Internet Archive. hans spemann. 1935 nobel Laureate in Medicine http://almaz.com/nobel/medicine/1935a.html

H ANS S PEMANN 1935 Nobel Laureate in Medicine for his discovery of the organizer effect in embryonic development. Background Residence: Germany Affiliation: University of Freiburg im Breisgau Featured Internet Links Search WWW Search The Nobel Prize Internet Archive Links added by Nobel Internet Archive visitors Back to The Nobel Prize Internet Archive Literature Peace Chemistry ... Medicine We always welcome your feedback and comments

4. Index Of Nobel Laureates In Medicine ALPHABETICAL LISTING OF nobel PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE. Name,Year Awarded. Snell, George D. 1980. spemann, hans, 1935. Sperry, Roger W. 1981. http://almaz.com/nobel/medicine/alpha.html

ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE Name Year Awarded Adrian, Lord Edgar Douglas Arber, Werner Axelrod, Julius Baltimore, David ... Medicine We always welcome your feedback and comments

5. Spemann, Hans spemann, hans. 12, 1941, Freiburg im Breisgau, Ger.), German embryologist who wasawarded the nobel Prize for Physiology or Medicine in 1935 for his discovery http://www.britannica.com/nobel/micro/560_31.html

Spemann, Hans Nobel Prize for Physiology or Medicine in 1935 for his discovery of the effect now known as embryonic induction, the influence exercised by various parts of the embryo that directs the development of groups of cells into particular tissues and organs. Spemann's concept of induction was based upon a lifetime of research into the early development of the newt. His work showed that, in the earliest stages, the fate of the embryonic parts has not been determined: if a piece of presumptive skin tissue is excised and transplanted into an area of presumptive nervous tissue, it will form nervous tissue, not skin. These results illuminated not only normal processes of development but also the origin of congenital abnormalities. Spemann summarized his researches in Embryonic Development and Induction).

6. Nobel Prize Winners For Physiology Or Medicine Whipple, George H. US, discoveries concerning liver treatment for anemia.1935, spemann, hans, Germany, organizer effect in embryo. 1936, http://www.britannica.com/nobel/table/phymed.html

Year Article Country* Achievement Behring, Emil von Germany work on serum therapy Ross, Sir Ronald U.K. discovery of how malaria enters an organism Finsen, Niels Ryberg Denmark treatment of skin diseases with light Pavlov, Ivan Petrovich Russia work on the physiology of digestion Koch, Robert Germany tuberculosis research Golgi, Camillo Italy work on the structure of the nervous system Spain work on the structure of the nervous system Laveran, Alphonse France discovery of the role of protozoa in diseases Ehrlich, Paul Germany work on immunity Russia work on immunity Kocher, Emil Theodor Switzerland physiology, pathology, and surgery of the thyroid gland Kossel, Albrecht Germany researches in cellular chemistry Gullstrand, Allvar Sweden work on dioptrics of the eye Carrel, Alexis France work on vascular suture; transplantation of organs Richet, Charles France work on anaphylaxis Austria-Hungary work on vestibular apparatus Bordet, Jules Belgium work on immunity factors in blood serum Krogh, August Denmark discovery of capillary motor-regulating mechanism Hill, A.V.

7. Spemann, Hans. The Columbia Encyclopedia, Sixth Edition. 2001 2001. spemann, hans. He received the 1935 nobel Prize in Physiology or Medicineand described his research in Embryonic Development and Induction (1938). http://www.bartleby.com/65/sp/Spemann.html

Select Search All Bartleby.com All Reference Columbia Encyclopedia World History Encyclopedia World Factbook Columbia Gazetteer American Heritage Coll. Dictionary Roget's Thesauri Roget's II: Thesaurus Roget's Int'l Thesaurus Quotations Bartlett's Quotations Columbia Quotations Simpson's Quotations English Usage Modern Usage American English Fowler's King's English Strunk's Style Mencken's Language Cambridge History The King James Bible Oxford Shakespeare Gray's Anatomy Farmer's Cookbook Post's Etiquette Bulfinch's Mythology Frazer's Golden Bough All Verse Anthologies Dickinson, E. Eliot, T.S. Frost, R. Hopkins, G.M. Keats, J. Lawrence, D.H. Masters, E.L. Sandburg, C. Sassoon, S. Whitman, W. Wordsworth, W. Yeats, W.B. All Nonfiction Harvard Classics American Essays Einstein's Relativity Grant, U.S. Roosevelt, T. Wells's History Presidential Inaugurals All Fiction Shelf of Fiction Ghost Stories Short Stories Shaw, G.B. Stein, G. Stevenson, R.L. Wells, H.G. Reference Columbia Encyclopedia PREVIOUS NEXT ... BIBLIOGRAPHIC RECORD The Columbia Encyclopedia, Sixth Edition. Spemann, Hans

8. Spemann, Hans. The American Heritage® Dictionary Of The English Language: Fourt 2000. spemann, hans. SYLLABICATION Spe·mann. PRONUNCIATION shp män. DATES 1869–1941.German zoologist and physiologist. He won a 1935 nobel Prize for his http://www.bartleby.com/61/50/S0625000.html

Select Search All Bartleby.com All Reference Columbia Encyclopedia World History Encyclopedia World Factbook Columbia Gazetteer American Heritage Coll. Dictionary Roget's Thesauri Roget's II: Thesaurus Roget's Int'l Thesaurus Quotations Bartlett's Quotations Columbia Quotations Simpson's Quotations English Usage Modern Usage American English Fowler's King's English Strunk's Style Mencken's Language Cambridge History The King James Bible Oxford Shakespeare Gray's Anatomy Farmer's Cookbook Post's Etiquette Bulfinch's Mythology Frazer's Golden Bough All Verse Anthologies Dickinson, E. Eliot, T.S. Frost, R. Hopkins, G.M. Keats, J. Lawrence, D.H. Masters, E.L. Sandburg, C. Sassoon, S. Whitman, W. Wordsworth, W. Yeats, W.B. All Nonfiction Harvard Classics American Essays Einstein's Relativity Grant, U.S. Roosevelt, T. Wells's History Presidential Inaugurals All Fiction Shelf of Fiction Ghost Stories Short Stories Shaw, G.B. Stein, G. Stevenson, R.L. Wells, H.G. Reference American Heritage Dictionary spelunker ... BIBLIOGRAPHIC RECORD The American Heritage Dictionary of the English Language: Fourth Edition. Spemann, Hans

9. Biography Of Hans Spemann 1919 Head of the Institute of Zoology, Freiburg; 1935 nobel price forMedicine; hans spemann died in Freiburg on September 12th, 1941. http://www.fml.tuebingen.mpg.de/spemann.htm

Hans Spemann (1869-1941) The biography : Hans Spemann was born in Stuttgart on June 27th, 1869. 1908 Professor in Rostock und Head of the Institute of Zoology, Rostock 1913 Head of the Department for Development and Causal Morphology of the newly founded 'Kaiser-Willhelm-Institute of Biology' in Berlin-Dahlem 1919 Head of the Institute of Zoology, Freiburg 1935 Nobel price for Medicine Hans Spemann died in Freiburg on September 12th, 1941 Hans Spemann studied embyogenesis on the newt embryo by transplantation and by cutting off the circulation to certain tissues. One of his most famous experiments together with H. Mangold was the transplantation of the blastopol lip in a Triturus embryo, wich led to the formation of a second body axsis, like in siamese twins. On vertebrate eyes, he discovered the induction of different Anlagen, for example the induction of the lens from epidermal tissue. For the first time he tried to find causal relations between different events of the embryogenesis. He was the first zoologist to recieve the Nobel price. Document last modified March 10th, 1998 by

10. Spemann, Hans spemann, hans , 1869–1941, German embryologist He received the 1935 nobel Prizein Physiology or Medicine and described his research in Embryonic Development http://www.infoplease.com/cgi-bin/id/A0846235

11. Nobel Prizes (table) encyclopediaEncyclopedia. nobel Prizes. 1935, Carl von Ossietzky, FrédéricJoliotCurie Irène Joliot-Curie, Sir James Chadwick, hans spemann, http://www.infoplease.com/ce6/sci/A0835783.html

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12. Scientist Profile : Hans Spemann hans spemann, a German embryologist, is one of the original pioneers of modern embryology,and one of only two embryologists to ever be awarded the nobel Prize http://hyperion.advanced.org/24355/data/details/profiles/spemann.html

Hans Spemann Hans Spemann Walter Sutton Oswald Avery Francis Crick James Watson ... Richard Seed Hans Spemann, a German embryologist, is one of the original pioneers of modern embryology , and one of only two embryologists to ever be awarded the Nobel Prize. His studies focused upon the differentiation of embryo cells during an organism's development. Spemann, born in Stuggart, Germany in 1869, attended the University of Heidelberg and the University of Munch before earning his Ph.D. in botany, zoology, and physics at the Zoological Institute of the University of Wurzberg in 1895. Spemann's fascination with embryology did not begin until after his college years, when during the winter of 1986 he read August Weismann's book The Germ Plasm: A Theory of Heredity as he sat isolated in a sanitarium, recovering from tuberculosis. In Spemann, ingeniously using a stand of hair as a noose, successfully split apart the

13. 1Up Info > Spemann, Hans (Cell Biology, Biographies) - Encyclopedia spemann, hanshäns shp ´män Pronunciation Key, 1869–1941, German embryologist Hereceived the 1935 nobel Prize in Physiology or Medicine and described his http://www.1upinfo.com/encyclopedia/S/Spemann.html

You are here 1Up Info Encyclopedia Cell Biology, Biographies Spemann, Hans ... News Search 1Up Info ENCYCLOPEDIA Cell Biology, Biographies Spemann, Hans Related Category: Cell Biology, Biographies Spemann, Hans Pronunciation Key Embryonic Development and Induction Read more articles on Spemann, Hans Click Here Related Resources and Utilities AMAZON All Products Books Magazines Popular Music Classical Music Video DVD Electronics Software Outdoor Living Wireless Phones Computers Outlet Read articles on eLibrary: In the Spotlight Severe Acute Respiratory Syndrome (SARS) Information Encyclopedia Topics History People Places Content on this web site is provided for informational purposes only. We accept no responsibility for any loss, injury or inconvenience sustained by any person resulting from information published on this site. We encourage you to verify any critical information with the relevant authorities. Home Contact Us Privacy Links Directory ©1Up Info

14. Zeal.com - United States - New - Library - Sciences - Chemistry - Disciplines - 1. spemann, hans 1935 nobel Biography http//www.nobel.se/medicine/laureates/1935/spemann-bio.htmlMeet the German scientist who discovered the organizer http://www.zeal.com/category/preview.jhtml?cid=331426

15. Spemann's Induction Experiments (Mangold died the year of her publication. nobel prizes are not given posthumously.). TheHeritage of Experimental Embryology hans spemann and the Organizer. http://zygote.swarthmore.edu/regul2.html

A Selective History of Induction II. Spemann's induction experiments A. Spemann's lens induction experiments In 1901, Curt Herbst wrote that he thought it possible "to establish the occurrence of formative stimuli which are exerted from one part of the embryo to another, and to demonstrate eventually the possiblity of a complete resolution of the entire ontogenesis into a sequence of such inductions." This prediction of inductive cascades was a bold statement, given that no sequence of events had yet been observed (1, 2). But that was soon to change. That same year, Hans Spemann (3) published "one of the most significant and seminal papers in the history of embryology" (4), his experimental analysis of lens formation in the frog. Spemann cauterized the prospective retina anlagen in the neurula stage Rana fusca Rana palustris (a related American frog) and got the opposite results: free lenses which were not connected to an eye. King's paper sent Spemann back to the laboratory where he did the identical experiment on different species of frog neurulae. He confirmed both his earlier results and Helen King's results (7). There was a component of induction that was species-specific. There appeared to be two ways to make a lens: induction or self-determination. Spemann put this result in the context of a model proposed the previous yearconcerning the operculum of amphibian tadpoles. When the limbs of amphibian tadpoles arise during metamorphosis, they emerge through an opening called the operculum. It had been thought that the limbs mechanically pushed through this barrier. However, when Herman Braus removed the forelimb rudiments, the operculum still openned at the appropriate time. Braus explained this phenomenon by introducing the engineering term "double assurance" into embryology (6). Spemann (8) maintained that double assurance also worked for amphibian lenses, and that while some frogs had only one mechanism of lens determination, most species were in between and had some self-differentiation but still required contact for optimum lens formation.

16. Premios Nobel De Medicina Premios nobel de Medicina. Año, Tema, Ganador. 1901, Behring, Emil Adolf Von.1902, Ross, Sir Ronald. 1935, spemann, hans. 1936, Dale, Sir Henry Hallett; Loewi,Otto. http://fai.unne.edu.ar/biologia/nobeles/nobelmed.htm

Premios Nobel de Medicina Tema Ganador Behring, Emil Adolf Von Ross, Sir Ronald Finsen, Niels Ryberg Pavlov, Ivan Petrovich Koch, Robert Cajal, Santiago Ramon Y.; Golgi, Camillo Laveran, Charles Louis Alphonse Ehrlich, Paul; Metchnikoff, Ilya Ilyich Kocher, Emil Theodor Kossel, Albrecht Gullstrand, Allvar Carrel, Alexis Richet, Charles Robert Barany, Robert Bordet, Jules Krogh, Schack August Steenberger Hill, Sir Archibald Vivian; Meyerhof, Otto Fritz; Banting, Sir Frederick Grant; Macleod, John James Richard; Einthoven, Willem; Fibiger, Johannes Andreas Grib Wagner-Jauregg, Julius Nicolle, Charles Jules Henri Eijkman, Christiaan; Hopkins, Sir Frederick Gowland Landsteiner, Karl Warburg, Otto Heinrich Adrian, Lord Edgar Douglas; Sherrington, Sir Charles Scott Morgan, Thomas Hunt Minot, George Richards; Murphy, William Parry; Whipple, George Hoyt Spemann, Hans Dale, Sir Henry Hallett; Loewi, Otto Nagyrapolt, Albert Szent-Gyorgyi Von Heymans, Corneille Jean Francois Domagk, Gerhard Dam, Henrik Carl Peter; Doisy, Edward Adelbert Erlanger, Joseph; Gasser, Herbert Spencer

17. ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN CHEMISTRY Zsigmondy, Richard Adolf, 1925. ALPHABETICAL LISTING OF nobel PRIZE LAUREATESIN PHYSIOLOGY AND MEDICINE. Snell, George D. 1980. spemann, hans, 1935. http://www.bioscience.org/urllists/nobelc.htm

FRONTIERS IN BIOSCIENCE; ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN CHEMISTRY, PHYSIOLOGY AND MEDICINE ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN CHEMISTRY Name Year Awarded Alder, Kurt Altman, Sidney Anfinsen, Christian B. Arrhenius, Svante August ... Zsigmondy, Richard Adolf ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE Name Year Awarded Adrian, Lord Edgar Douglas Arber, Werner Axelrod, Julius Baltimore, David ... Zinkernagel, Rolf M. Source: The Nobel Prize Internet Archive

18. ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE FRONTIERS IN BIOSCIENCE; ALPHABETICAL LISTING OF nobel PRIZE LAUREATES IN PHYSIOLOGYAND MEDICINE, Name, Year Awarded. Snell, George D. 1980. spemann, hans, 1935. http://www.bioscience.org/urllists/nobelm.htm

FRONTIERS IN BIOSCIENCE; ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE Name Year Awarded Adrian, Lord Edgar Douglas Arber, Werner Axelrod, Julius Baltimore, David ... Zinkernagel, Rolf M.

19. Spemann, Hans spemann, hans 18691941, German embryologist He received the 1935 nobel Prize inPhysiology or Medicine and described his research in Embryonic Development and http://www.slider.com/enc/50000/Spemann_Hans.htm

energy drinks Home Encyclopeadia S Sou - Spe ... Rope Ladders Trellian WebPage Slider Search: The Web Encyclopaedia Shopping Index Help Encyclopaedia Spemann, Hans 1869-1941, German embryologist. He was professor of zoology (1919-35) at the Univ. of Freiburg. By transplanting embryonic tissue to a new location or to another embryo, he investigated the agency that governs the growth and differentiation of cells. He received the 1935 Nobel Prize in Physiology or Medicine and described his research in Embryonic Development and Induction Add URL Advertise Contact Us

20. Spemann, Hans Translate this page de Freiburg-im-Breisgau, dans la succession à hans Doflein, un Le nom de Spemannsera toujours associé à son travail on lui a attribué le Prix nobel en 1935 http://www.cartage.org.lb/fr/themes/Biographies/mainbiographie/S/Spemann/Spemann

Spemann, Hans Hans Spemann est né le 27 juin 1869, à Stuttgart. Il était le fils l'aîné de l'éditeur, Wilhelm Spemann. De 1878 jusqu'à 1888 il est allé à l'Ecole Eberhard-Ludwig à Stuttgart et quand il a laissé(quitté) l'école en 1888 il a dépensé(passé) une année dans l'affaire de publication de son père. De 1889-1890 il a fait son service militaire et ensuite, après une période comme un libraire au détail, il est entré, en 1891, l'Université de Heidelberg. Là, avant qu'il n'ait pris son examen préliminaire en 1893, il a étudié la médecine et a été particulièrement attiré par le travail de l'anatomiste comparatif là, Carl Gegenbaur. Pendant l'hiver de 1893-1894 il a étudié à l'Université de Munich, où il lui est devenu plus étroitement(de près) mis au courant août Pauly - un fait très important. Du printemps de 1894 à la fin de 1908, il a travaillé dans l'Institut Zoologique à l'Université de Würzburg. En 1895 il a pris son degré dans la zoologie, la botanique et la physique (des sujets pour servir ses études anatomiques), ayant travaillé sous Theodor Boveri, Julius Sachs et Wilhelm RoNtgen, tout de ce qui avait l'influence la plus grande sur son développement scientifique. En 1898 il a qualifié comme un conférencier dans la zoologie à l'Université de Würzburg et en 1908 on lui a demandé de devenir le Professeur de Zoologie et l'Anatomie Comparative à Rostock et en 1914 il est devenu le Directeur d'Associé du Kaiser Wilhelm l'Institut de Biologie à Berlin-Dahlem. En 1919 il a été nommé le Professeur de Zoologie à l'Université de Freiburg-im-Breisgau, dans la succession à Hans Doflein, un poste qu'il s'est tenu avant qu'il n'ait mis à la retraite et soit devenu le Professeur honoraire en 1935.

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