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         Hartline Haldan Keffer:     more detail

1. Haldan Keffer Hartline Winner Of The 1967 Nobel Prize In Medicine
haldan keffer hartline, a nobel Prize Laureate in Physiology and Medicine,at the nobel Prize Internet Archive. haldan keffer hartline.
http://almaz.com/nobel/medicine/1967b.html
H ALDAN K EFFER H ARTLINE
1967 Nobel Laureate in Medicine
    for their discoveries concerning the primary physiological and chemical visual processes in the eye.
Background

    Residence: U.S.A.
    Affiliation: The Rockefeller University, New York, NY
Featured Internet Links Links added by Nobel Internet Archive visitors Back to The Nobel Prize Internet Archive
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2. Haldan K. Hartline - Biography
In 1936 haldan keffer hartline married Elizabeth Kraus, daughter of the eminentchemist CA Kraus. Daniel keffer and Peter haldan have positions in
http://www.nobel.se/medicine/laureates/1967/hartline-bio.html
Haldan Keffer Hartline was born in Bloomsburg, Pennsylvania, on December 22nd, 1903. His parents were teachers there in the State Normal School (now Bloomsburg State College) where he received his early education. His father, Daniel S. Hartline, was Professor of Biology, but a man whose wide interests also included Astronomy and Geology. It was through his father that Keffer became interested in Natural Sciences.
Keffer Hartline attended Lafayette College in Easton, Pennsylvania, graduating in 1923 (B. Sc.). His college teacher of biology, Beverly W. Kunkel, encouraged him to undertake research; his first scientific paper concerned visual responses of land isopods. Summers at the Marine Biological Laboratory at Woods Hole added to his biological training; there he was especially influenced by Jacques Loeb, Selig Hecht, and Merkel H. Jacobs.
In the autumn of 1923 he entered the Johns Hopkins School where he was encouraged to continue his research interest in vision in the Department of Physiology under E. K. Marshall and C. D. Snyder. Dr. Snyder let him use his Einthoven string galvanometer with which Hartline undertook the study of the retinal action potential using frogs, decerebrate cats and rabbits. He learned to obtain electroretinograms from intact animals, and recorded clearly recognizable retinal action potentials from human subjects. He also used intact insects for quantitative studies.

3. Hartline, Haldan Keffer
hartline, haldan keffer. 17, 1983, Fallston, Md.), American physiologist who wasa cowinner (with George Wald and Ragnar Granit) of the 1967 nobel Prize in
http://www.britannica.com/nobel/micro/260_49.html
Hartline, Haldan Keffer
(b. Dec. 22, 1903, Bloomsburg, Pa., U.S.d. March 17, 1983, Fallston, Md.), American physiologist who was a cowinner (with George Wald and Ragnar Granit ) of the 1967 Nobel Prize in Physiology or Medicine for his work in analyzing the neurophysiological mechanisms of vision. Hartline began his study of retinal electrophysiology as a National Research Council Fellow at Johns Hopkins University, Baltimore, receiving his M.D. in 1927. After attending the universities of Leipzig and Munich as an Eldridge Johnson traveling research scholar, he became professor of biophysics and chairman of the department at Johns Hopkins in 1949. He joined the staff of Rockefeller University, New York City, in 1953 as professor of neurophysiology. Hartline investigated the electrical responses of the retinas of certain arthropods, vertebrates, and mollusks because their visual systems are much simpler than those of humans and are thus easier to study. He concentrated his studies on the eye of the horseshoe crab ( Limulus polyphemus ). Using minute electrodes in his experiments, he obtained the first record of the electrical impulses sent by a single optic nerve

4. Search Results For Haldan Keffer Hartline - Encyclopædia Britannica - The Onlin
haldan keffer hartline nobel Foundation Biography of this American scientist awardedthe nobel Prize in Medicine or Physiology, in 1967, for research on the
http://www.britannica.com/search?query=haldan keffer hartline&seo

5. Nat'l Academies Press, Biographical Memoirs (1990), Haldan Keffer Hartline
OCR for page 213 haldan keffer hartline 213 1969 Visual receptors and retinal interaction.In Les Prix nobel en 1967, The nobel Foundation, Stockholm, pp.242
http://www.nap.edu/books/0309041988/html/196.html
Biographical Memoirs V.59
National Academy of Sciences ( NAS
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Openbook Linked Table of Contents Front Matter, pp. i-iv Table of Contents, pp. v-vi Preface, pp. vii-viii Title Page, pp. 1-1 Frederic C. Bartter, pp. 2-25 George Wells Beadle, pp. 26-53 Solomon A. Berson, pp. 54-71 Raymond Thayer Birge, pp. 72-85 William Henry Chandler, pp. 86-115 Gertrude Mary Cox, pp. 116-133 Conrad Arnold Elvehjem, pp. 134-167 Gottfried Samuel Fraenkel, pp. 168-195 Haldan Keffer Hartline, pp. 196-213 Mark Kac, pp. 214-235 Aldo Starker Leopold, pp. 236-255 Manfred Martin Mayer, pp. 256-281 Walsh McDermott, pp. 282-307 Theophilus Shickel Painter, pp. 308-337 Edward Lawrie Tatum, pp. 356-387 Cornelis Bernardus Van Niel, pp. 388-423 Robert H. Whittaker, pp. 424-445 Maxwell Mayer Wintrobe, pp. 446-472 Cumulative Index, pp. 473-486
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6. Themes Geography History History Prize Winners Nobel
Themes Geography History History Prize Winners nobel Prize Medicine.Year, Winners. 1967, Granit, Ragnar hartline, haldan keffer - Wald, George.
http://www.cartage.org.lb/en/themes/GeogHist/histories/prizewinners/nobelprize/m
Themes History Prize Winners Nobel Prize
Year
Winners
Behring, Emil Adolf von Ross, Ronald Finsen, Niels Ryberg Pavlov, Ivan Petrovich ... Bárány, Robert The prize money was allocated to the Special Fund of this prize section The prize money was allocated to the Special Fund of this prize section The prize money was allocated to the Special Fund of this prize section The prize money was allocated to the Special Fund of this prize section Bordet, Jules Krogh, Schack August Steenberg The prize money was allocated to the Special Fund of this prize section Hill, Archibald Vivian Meyerhof, Otto Fritz Banting, Frederick Grant Macleod, John James Richard ... Einthoven, Willem The prize money was allocated to the Special Fund of this prize section Fibiger, Johannes Andreas Grib Wagner-Jauregg, Julius Nicolle, Charles Jules Henri Eijkman, Christiaan ... Domagk, Gerhard The prize money was 1/3 allocated to the Main Fund and 2/3 to the Special Fund of this prize section The prize money was 1/3 allocated to the Main Fund and 2/3 to the Special Fund of this prize section The prize money was 1/3 allocated to the Main Fund and 2/3 to the Special Fund of this prize section Dam, Henrik Carl Peter

7. NASA Neurolab Web: Mission Home Page
haldan keffer hartline (19031983), American biophysicist, first mechanisms of vision,hartline's research contributed to He shared the 1967 nobel Prize in
http://neurolab.jsc.nasa.gov/hartline.htm
Spotlight on Neuroscience
Haldan Keffer Hartline
Haldan Keffer Hartline (1903-1983), American biophysicist, first determined the mechanics of how the nerves of the visual system receive information and transfer it to the brain. He was born in Bloomsburg, Pennsylvania. Hartline worked with horseshoe crabs and frogs to investigate the mechanisms of vision. His discoveries convinced researchers that the retina (the innermost layer of the eye that is light sensitive) and optic nerves themselves process many nerve signals before the signals are transmitted to the brain. Besides providing new knowledge about the neurophysiological mechanisms of vision, Hartline's research contributed to advances in night-vision, pattern-recognition, and motion-detection devices. He shared the 1967 Nobel Prize in physiology or medicine with Swedish neurophysiologist Ragnar Granit and American biochemist George Wald for their work on the biochemical and electrical processes of vision. Curator: Julie Heath and Jacque Havelka
Responsible NASA Official: Katherine Newkirk
Updated 9 March 1998

8. Premios Nobel De Medicina
Premios nobel de Medicina. Año, Tema, Ganador. 1966, Huggins, Charles Brenton;Rous, Peyton. 1967, Granit, Ragnar; hartline, haldan keffer; Wald, George.
http://fai.unne.edu.ar/biologia/nobeles/nobelmed.htm
Premios Nobel de Medicina
Tema Ganador Behring, Emil Adolf Von Ross, Sir Ronald Finsen, Niels Ryberg Pavlov, Ivan Petrovich Koch, Robert Cajal, Santiago Ramon Y.; Golgi, Camillo Laveran, Charles Louis Alphonse Ehrlich, Paul; Metchnikoff, Ilya Ilyich Kocher, Emil Theodor Kossel, Albrecht Gullstrand, Allvar Carrel, Alexis Richet, Charles Robert Barany, Robert Bordet, Jules Krogh, Schack August Steenberger Hill, Sir Archibald Vivian; Meyerhof, Otto Fritz; Banting, Sir Frederick Grant; Macleod, John James Richard; Einthoven, Willem; Fibiger, Johannes Andreas Grib Wagner-Jauregg, Julius Nicolle, Charles Jules Henri Eijkman, Christiaan; Hopkins, Sir Frederick Gowland Landsteiner, Karl Warburg, Otto Heinrich Adrian, Lord Edgar Douglas; Sherrington, Sir Charles Scott Morgan, Thomas Hunt Minot, George Richards; Murphy, William Parry; Whipple, George Hoyt Spemann, Hans Dale, Sir Henry Hallett; Loewi, Otto Nagyrapolt, Albert Szent-Gyorgyi Von Heymans, Corneille Jean Francois Domagk, Gerhard Dam, Henrik Carl Peter; Doisy, Edward Adelbert Erlanger, Joseph; Gasser, Herbert Spencer

9. Haldan Keffer Hartline: Awards Won By Haldan Keffer Hartline
123Awards hardwork is paid in form of awards. Awards of haldan keffer hartline.OTHERnobel, 1967, MEDICINE. Enter Artist/Album. Partner Sites. Stardose.com.
http://www.123awards.com/artist/1463.asp
hardwork is paid in form of awards Awards of Haldan Keffer Hartline OTHER-NOBEL MEDICINE Enter Artist/Album
Partner Sites
Stardose.com RealLyrics.com OnlyHitLyrics.com Biography Search Engine ... privacy

10. Search Results For Sandbæk Ragnar - Encyclopædia Britannica - The Online Enc
hartline, haldan keffer American physiologist who was a cowinner (with George Waldand Ragnar Granit) of the 1967 nobel Prize for Physiology or Medicine for
http://search.britannica.com/search?ct=eb&query=Sandbæk Ragnar

11. ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND
FRONTIERS IN BIOSCIENCE; ALPHABETICAL LISTING OF nobel PRIZE LAUREATES INPHYSIOLOGY AND MEDICINE, Name, Year Awarded. hartline, haldan keffer, 1967.
http://www.bioscience.org/urllists/nobelm.htm

12. ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN CHEMISTRY
Zsigmondy, Richard Adolf, 1925. ALPHABETICAL LISTING OF nobel PRIZE LAUREATES INPHYSIOLOGY AND MEDICINE. Gullstrand, Allvar, 1911. hartline, haldan keffer, 1967.
http://www.bioscience.org/urllists/nobelc.htm
FRONTIERS IN BIOSCIENCE;
ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN
CHEMISTRY, PHYSIOLOGY AND MEDICINE

ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN CHEMISTRY Name Year Awarded Alder, Kurt Altman, Sidney Anfinsen, Christian B. Arrhenius, Svante August ... Zsigmondy, Richard Adolf ALPHABETICAL LISTING OF NOBEL PRIZE LAUREATES IN PHYSIOLOGY AND MEDICINE Name Year Awarded Adrian, Lord Edgar Douglas Arber, Werner Axelrod, Julius Baltimore, David ... Zinkernagel, Rolf M. Source: The Nobel Prize Internet Archive

13. Sigma Xi: The Scientific Research Society: Nobel Laureates
About Sigma Xi » Overview » nobel Laureates 1964 Konrad E. Bloch 1966 CharlesB. Huggins 1966 Francis Peyton Rous 1967 haldan keffer hartline 1967 George
http://www.sigmaxi.org/about/overview/nobel.shtml
Overview Leadership Organization News ... Contact Us About: Overview
Overview
Physics
1907 Albert Michelson
1921 Albert Einstein
1923 Robert A. Millikan
1925 James Franck
1927 Arthur H. Compton
1936 Carl D. Anderson
1937 Clinton J. Davisson 1938 Enrico Fermi 1939 Ernest O. Lawrence 1943 Otto Stern 1944 Isidor Isaac Rabi 1945 Wolfgang Pauli 1946 Percy Williams Bridgman 1952 Felix Bloch 1952 Edward M. Purcell 1955 Polykarp Kusch 1955 Willis E. Lamb, Jr. 1956 John Bardeen 1956 Walter H. Brattain 1956 William Shockley 1957 Chen Ning Yang 1958 Igor Y. Tamm 1959 Owen Chamberlain 1959 Emilio G. Segre 1960 Donald A. Glaser 1961 Robert Hofstadter 1963 Eugene P. Wigner

14. The Johns Hopkins:nobel Prize Winners
Medicine, 1966. haldan keffer hartline MD 1927; Professor of Biophysics,194954 nobel Prize in Medicine, 1967. Lars Onsager Associate
http://webapps.jhu.edu/jhuniverse/information_about_hopkins/facts_and_statistics

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nobel prize winners
Woodrow Wilson
Ph.D. 1886 (History)
Nobel Prize in Peace, 1919 James Franck
Professor of Physics, 1935-38
Nobel Prize in Physics, 1925 Nicholas Murray Butler
Lecturer, 1890-91
Nobel Prize in Peace, 1931 Thomas Hunt Morgan Ph.D. 1890 (Zoology); LL.D. 1915 Nobel Prize in Medicine, 1933 George Richards Minot Assistant in Medicine, 1914-15 Nobel Prize in Medicine, 1934 George Hoyt Whipple M.D. 1905; Associate Professor in Pathology, 1910-14 Nobel Prize in Medicine, 1934 Harold Clayton Urey Associate in Chemistry, 1924-28 Nobel Prize in Chemistry, 1934 Joseph Erlanger M.D. 1899; Assistant in Physiology, 1900-01; Instructor, 1901-03; Associate, 1903-04; Associate Professor, 1904-06; LL.D. 1947 Nobel Prize in Medicine, 1944 Herbert Spencer Gasser M.D. 1915 Nobel Prize in Physiology, 1944 Vincent du Vigneaud National Research Fellow, Pharmacology 1927-28 Nobel Prize in Chemistry, 1955

15. Physiology Or Medicine 1984
1967 The prize was awarded jointly to GRANIT, RAGNAR, hartline, haldan keffer,. 1967A díjat megosztva kapták GRANIT, RAGNAR, hartline, haldan keffer,.
http://www.radnoti.hu/common/nobel/orvos.htm
The prize was awarded jointly to: JERNE , NIELS K.,
KÖHLER , GEORGES J.F., and MILSTEIN , CÉSAR, (photo)
Great Britain and Argentina, MRC Laboratory of Molecular Biology, Cambridge,
b. 1927 (in Bahia Blanca, Argentina)
"for theories concerning the specificity in development and control of the immune system and the discovery of the principle for production of monoclonal antibodies" A díjat megoszva kapták: JERNE , NIELS K.,
KÖHLER , GEORGES J.F., és MILSTEIN , CÉSAR, (kép)
Great Britain and Argentina, MRC Laboratory of Molecular Biology, Cambridge,
“az immunrendszer fejlõdésében és szabályozásában megnyilvánuló specificitásra vonatkozó elméletekért és a monoklonális antitestek képzõdési elvének felfedezéséért” The prize was awarded jointly to: BERGSTRÖM , SUNE K.,
SAMUELSSON , BENGT I., and VANE , Sir JOHN, (photo)
Great Britain, The Wellcome Research Laboratories, Beckenham,
b. 1927 "for their discoveries concerning prostaglandins and related biologically active substances" A díjat megosztva kapták: BERGSTRÖM , SUNE K.

16. University Of Chicago News: Nobel Laureates
The nobel Prize in Physiology or Medicine 1967 with haldan keffer hartline and RagnarGranit “for their discoveries concerning the primary physiological and
http://www-news.uchicago.edu/resources/nobel/physmed.html
University of Chicago News: Resources
University of Chicago Physiology or Medicine Nobel Laureates Seventy-four Laureates have been faculty, students or researchers at the University of Chicago. Eleven of those Laureates won prizes in Physiology or Medicine.
Roger W. Sperry

Ph.D., 1941; Assistant Professor in the Department of Anatomy , 1946-53; Assistant Professor in the Departments of Anatomy and Psychology The Nobel Prize in Physiology or Medicine 1981
with David H. Hubel, M.D., and Torsten N. Wiesel, M.D.
George Wald

Postdoctoral Fellow, 1932-34. The Nobel Prize in Physiology or Medicine 1967
with Haldan Keffer Hartline and Ragnar Granit
Charles Brenton Huggins

Instructor of Surgery, 1927-29; Assistant Professor, 1929-33; Associate Professor, 1933-36; Professor, 1936; Director of the Ben May Laboratory for Cancer Research , 1951-69; William B. Ogden Distinguished Service Professor, 1962-present. The Nobel Prize in Physiology or Medicine 1966
with Peyton Rous Konrad Bloch Assistant Professor in the Institute of Radiobiology and in the Departments of Biochemistry and Biophysics, 1946-48; Associate Professor, 1948-54; Professor, 1954. The Nobel Prize in Physiology or Medicine 1964 with Feodor Lynen Sir John Carew Eccles Professorial Lecturer in the Department of Physiology The Nobel Prize in Physiology or Medicine 1963 with Sir Alan Lloyd Hodgkin and Sir Andrew Fielding Huxley James Dewey Watson Ph.B., 1946; S.B., 1947; D.Sc. (honorary), 1961.

17. Nobel Prize In Physiology Or Medicine - Wikipedia
Source http//www.nobel.se/medicine/laureates/index.html. 1966 Peyton Rous, CharlesBrenton Huggins 1967 Ragnar Granit, haldan keffer hartline, George Wald
http://www.wikipedia.org/wiki/Nobel_Prize/Physiology_or_medicine
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Nobel Prize in Physiology or Medicine
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Emil Adolf von Behring Ronald Ross Niels Ryberg Finsen ... Christiaan Eijkman , Sir Frederick Gowland Hopkins Karl Landsteiner Otto Heinrich Warburg Sir Charles Scott Sherrington Edgar Douglas Adrian Thomas Hunt Morgan George Hoyt Whipple ... Hans Spemann Sir Henry Hallett Dale Otto Loewi Albert von Szent-Györgyi Nagyrapolt Corneille Jean François Heymans ... Herbert Spencer Gasser Sir Alexander Fleming Ernst Boris Chain , Sir Howard Walter Florey Hermann Joseph Muller Carl Ferdinand Cori Gerty Theresa , née Radnitz Cori, Bernardo Alberto Houssay Paul Hermann Müller Walter Rudolf Hess Antonio Caetano De Abreu Freire Egas Moniz ... Dickinson W. Richards

18. Nobel Prize In Medicine Since 1901
Rous, Peyton. 1967, Granit, Ragnar; hartline, haldan keffer; Wald, George.
http://www.planet101.com/nobel_medi_hist.htm
Nobel Prize in Medicine since 1901 Year Prize Winners Behring, Emil Adolf Von Ross, Sir Ronald Finsen, Niels Ryberg Pavlov, Ivan Petrovich Koch, Robert Cajal, Santiago Ramon Y.; Golgi, Camillo Laveran, Charles Louis Alphonse Ehrlich, Paul; Mechnikov, Ilya Ilyich Kocher, Emil Theodor Kossel, Albrecht Gullstrand, Allvar Carrel, Alexis Richet, Charles Robert Barany, Robert Bordet, Jules Krogh, Schack August Steenberger Hill, Sir Archibald Vivian; Meyerhof, Otto Fritz; Banting, Sir Frederick Grant; Macleod, John James Richard; Einthoven, Willem; Fibiger, Johannes Andreas Grib Wagner-Jauregg, Julius Nicolle, Charles Jules Henri Eijkman, Christiaan; Hopkins, Sir Frederick Gowland Landsteiner, Karl Warburg, Otto Heinrich Adrian, Lord Edgar Douglas; Sherrington, Sir Charles Scott Morgan, Thomas Hunt Minot, George Richards; Murphy, William Parry; Whipple, George Hoyt Spemann, Hans Dale, Sir Henry Hallett; Loewi, Otto Nagyrapolt, Albert Szent-Gyorgyi Von Heymans, Corneille Jean Francois Domagk, Gerhard Dam, Henrik Carl Peter; Doisy, Edward Adelbert Erlanger, Joseph; Gasser, Herbert Spencer

19. The Nobel Prize
Neuroscience nobel Laureats The nobel Prize has been awarded many times to Neuroscientists 1967 USA. haldan keffer hartline. Mechanisms of vision. 1967 - USA.
http://pdbio.byu.edu/neuroscience/pages/nobel_prize.htm
Neuroscience Nobel Laureats: The Nobel Prize has been awarded many times to Neuroscientists: 1997 - U.S.A Stanley B. Prusiner Discovery of Prions; a new biological principle of infection 1994 - U.S.A. Alfred G. Gilman Discovery of G-Protein coupled receptors and their role in signal transduction 1994 - U.S.A. Martin Rodbell Discovery of G-Protein coupled receptors and their role in signal transduction 1991 - Germany Erwin Neher Function of single ion channels in cells 1991 - Germany Bert Sakmann Function of single ion channels in cells 1986 - U.S.A. Stanley Cohen Control of nerve cell growth 1986 - Italian U.S.A. Rita Levi-Montalcini Control of nerve cell growth 1982 - Sewden Sune K. D. Bergström Discovery of prostaglandinds 1982 - Sweden Bengt I. Samuelsson Discovery of prostaglandinds 1982 - Britain John Robert Vane Discovery of Prostaglandins 1981 - Swedish - U.S.A. Torsten N. Wiesel I nformation processing in the visual system 1981 - U.S.A. Roger W. Sperry Functions of the right and left hemispheres of the brain 1981 - Candian U.S.A.

20. SIMR - Centenary Survey Of Nobel Laureates
case in many places. Michael S Brown, MD, nobel Prizewinner 1985. 1967 Ragnar GRANIT,haldan keffer hartline and George WALD - describe the physiological and
http://www.simr.org.uk/pages/nobel/time_line_7.html
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Nobel Prize Winners in Physiology and Medicine (1960 - 1969)
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"I agree that animal experimentation should be humane and regulated, but the regulations must not be so onerous as to impede legitimate experimentation, which is now the case in many places." - Michael S Brown, M.D., Nobel Prizewinner 1985 Sir Frank MacFarlane BURNET and Sir Peter Brian MEDAWAR - describe acquired immunological tolerance. Georg von BŠKŠSY - shows how the inner ear works. Francis Harry Comptom CRICK, James Dewey WATSON and Maurice Hugh Frederick WILKINS - discover the molecular structure of DNA and RNA and how they transmit genetic information. Sir John Carew ECCLES, Sir Alan Lloyd HODGKIN and Sir Andrew Fielding HUXLEY - discover the ionic mechanisms by which nerves function. Peyton ROUS - discovers tumour-inducing viruses. Charles Brenton HUGGINS - devises the hormonal treatment of prostatic cancer.

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